Regarding pulmonary perfusion, which statement is INCORRECT?

• A. apical perfusion is less than basal
• B. E.coli endotoxin causes venodilatation
• C. total pulmonary blood flow increases 3-6 fold during exercise
• D. pulmonary vascular resistance increases at small lung volumes
• E. in zone 2, pulmonary arterial pressure is greater than alveolar pressure