If ‘A + B’ means ‘A is the son of B’; ‘A - B’ means ‘A is the daughter of B’, ‘A & B’ means ‘A is the father of B’ and ‘A @ B’ means ‘A is the mother of B’, then what does P + R & Q - S mean?

• A. R is the daughter of Q
• B. P is the sister of Q
• C. S is the brother of Q
• D. S is the mother of P